I feel like we are saying different things. When you manually cycled the bolt, did the rifle remove the rounds from the chamber every time?Both are issues present. When being fired, it was picked up 4/5 times, just not enough pressure to fully cycle the bolt. I have no idea why that changed when manually cycling?
Granted the velocity and forces involved under recoil are different but if the rifle isn't grabbing the rounds under manual cycling, it won't under recoil, either...and vice versa.